Clojure protocol error only during CI

I’m getting a “No implementation of method” exception with a clojure protocol, that only happens during the CI run of ‘lein trampoline test’. The test runs fine locally, and runs fine on the circleci VM when I ssh in and manually run ‘lein trampoline test’, or ‘lein test’ from the shell.

It only fails in the CI build. Is there something different about the build environment, vs. ssh, that would affect clojure/jvm?

Build log here:
https://circleci.com/gh/ucscXena/ucsc-xena-server/2